Hierophis viridiflavus carbonarius

Combinations

• Coluber viridiflavus var. carbonarius Bonaparte, 1833: unnumbered [original combination]
• Haemorrhois [sic] viridiflavus carbonarius — Welch, 1983 [subsequent combination]
• Hierophis viridiflavus carbonarius — Utiger & Schätti, 2004 [subsequent combination]
• Hierophis carbonarius — Mezzasalma, Dall'Asta, Loy, Cheylan, Lymberakis, Zuffi, Tomovic, Odierna, Guarino, 2015 [subsequent combination]

Synonyms

• Coluber gemonensis gyarosensis Mertens, 1968: 181 [synonymised by Utiger & Schätti (2004: 235)]
  • Holotype: ZFMK 4332, ♂, collected by K.F. Buchholz on 28-31 August 1963.
  • Type locality: “Insel Gyaros (Gioura), Cycladen”.

Primary references

• Bonaparte, C.L. 1832–1841. Iconographia della Fauna Italica per le Quattro Classi degli Animali Vertebrati. Tomo II. Amphibi. Salviucci: Rome. 65 pp. + 153 pls. BHL Reference page. 
• Mertens, R. 1968. Eine schwarze Zornnatter von den Cycladen: Coluber gemonensis gyarosensis n. subsp.. Senckenbergiana biologica 49(3–4): 181–189.
• Utiger, U. & Schätti, B. 2004. Morphology and phylogenetic relationships of the Cyprus racer, Hierophis cypriensis, and the systematic status of Coluber gemonensis gyarosensis Mertens (Reptilia: Squamata: Colubrinae). Revue Suisse de Zoologie 111(1): 225–238. BHL Reference page. 

Additional references

• Schätti, B. & Vanni, S. 1986. Intraspecific Variation in Coluber viridiflavus Lacépède, 1789, and validity of its subspecies (Reptilia, Serpentes, Colubridae). Revue suisse de Zoologie 93(1): 219–232. BHL Reference page. 
• Nagy, Z.T., Joger, U., Guicking, D. & Wink, M. 2002. Phylogeography of the European Whip Snake, Coluber (Hierophis) viridiflavus as inferred from nucleotide sequences of the mitochondrial cytochrome b gene and ISSR genomic fingerprinting. Biota 3(1–2): 109–118. ResearchGate Reference page. 

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• Mezzasalma, M., Dall’Asta, A., Loy, A., Cheylan, M., Lymberakis, P., Zuffi, M.A.L., Tomović, L., Odierna, G. & Guarino, F.M. 2015. A sisters’ story: a multidisciplinary study on the comparative phylogeography and taxonomy of Hierophis viridiflavus and H. gemonensis (Serpentes, Colubridae). Zoologica Scripta 44(5): 495–508. DOI: 10.1111/zsc.12115 Reference page. 

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• Speybroeck, J., Beukema, W., Dufresnes, C., Fritz, U., Jablonski, D., Lymberakis, P., Martínez-Solano, I., Razzetti, E., Vamberger, M., Vences, M., Vörös, J. & Crochet, P.-A. 2020. Species list of the European herpetofauna – 2020 update by the Taxonomic Committee of the Societas Europaea Herpetologica. Amphibia-Reptilia 41(2): 139–189. DOI: 10.1163/15685381-bja10010 Reference page. 
• Boundy, J. 2021. Snakes of the World: A Supplement. CRC Press, Taylor and Francis Group: Boca Raton, Florida. 273 pp. ISBN 9781138618138 ebook Reference page. 

Links

• Uetz, P., Freed, P., Aguilar, P., Reyes, R., Kudera, F. & Hošek, J. (eds.) 2025. Hierophis viridiflavus. The Reptile Database. Accessed on 11 June 2023.