Notocupes

Notocupes alienus (A–B), Notocupes minusculus (C–F)

Taxonavigation

Taxonavigation: Archostemata
Superregnum: Eukaryota Cladus: Unikonta Cladus: Opisthokonta Cladus: Holozoa Regnum: Animalia Subregnum: Eumetazoa Cladus: ParaHoxozoa Cladus: Bilateria Cladus: Nephrozoa Cladus: Protostomia Cladus: Ecdysozoa Cladus: Panarthropoda Phylum: Arthropoda Cladus: Pancrustacea Cladus: Allotriocarida Subphylum: Hexapoda Classis: Insecta Cladus: Dicondylia Subclassis: Pterygota Infraclassis: Neoptera Cladus: Eumetabola Cladus: Holometabola Superordo: Coleopterida Ordo: Coleoptera Subordo: Archostemata

Familia: incertae sedis
Genus: †Notocupes
Species (after Strelnikova & Yan (2021, 2023a, 2023b); Li et al. (2023a, 2023b)): †N. alienus – †N. baojiatunensis – †N. caudatus – †N. cyclodontus – †N. daohugouensis – †N. diazromerali – †N. elegans – †N. excellens – †N. kezuoensis – †N. khasurtyiensis – †N. laetus – †N. laiyangensis – (?)†N. lapidarius – †N. latus – †N. lini – †N. ludongensis – †N. mongolicus – †N. neli – †N. ohmkuhnlei – †N. picturatus – †N. pingi – (?)†N. pulcher – †N. reticulatus – †N. robustus – †N. rudis – †N. shiluoensis – †N. tuanwangensis – †N. validus

Transferred to Rhabdocupes (Strelnikova & Yan (2023b)): †N. celluosus – †N. dischdes – †N. issykkulensis – †N. laticella – †N. lentus – †N. minusculus – †N. oxypygus – †N. protensus – †N. rostratus – †N. sogutensis – †N. tenuis – †N. vitimensis

Transferred to Conexicoxa (Strelnikova & Yan (2023b)): †N. brachycephalus – †N. crassus – †N. denticollis – †N. epicharis – †N. eumeurus – †N. exiguus – †N. homorus – †N. jurassicus – †N. khetanensis – †N. kirghizicus – †N. longicollis – †N. martinclosas – †N. porrectus – †N. psilatus – †N. siniestri – †N. stabilis – †N. tripartitus

Transferred to Brachilatus (Strelnikova & Yan (2023b)): †N. caducus – †N. dundulaensis – †N. foersteri – †N. longicoxa – †N. nigrimonticola – †N. oculatus – †N. premeris – †N. spinosus – †N. viridis

Transferred to Odontomma (Strelnikova & Yan (2023b)): †N. patulus

Name

Notocupes Ponomarenko, 1964: 61

Type species: Notocupes picturatus Ponomarenko, 1964, by monotypy.

Synonyms

Sinocupes Lin, 1976: 113 [syn. Kirejtshuk et al. (2010: 788)]
- Type species: Sinocupes validus Lin, 1976, by monotypy.
Chengdecupes Hong, 1983: 83
- Type species: Chengdecupes jurassicus Hong, 1983, by original designation [and monotypy].
?Conexicoxa Lin, 1986: 76
- Type species: Conexicoxa homora Lin, 1986, by monotypy.
?Forticupes Hong & Wang, 1990: 105
- Type species: Forticupes laiyangensis Hong & Wang, 1990, by original designation [and monotypy].
?Picticupes Hong & Wang, 1990: 107
- Type species: Picticupes tuanwangensis Hong & Wang, 1990, by original designation [and monotypy].
Amblomma Tan, Ren & Liu, 2005: 208 [syn. Kirejtshuk et al. (2010: 788)]
- Type species: Amblomma psilata Tan, Ren & Liu, 2005, by original designation.
Ovatocupes Tan & Ren, 2006: 225 [syn. Kirejtshuk et al. (2010: 788)]
- Type species: Ovatocupes alienus Tan & Ren, 2006, by monotypy.
Echinocups Kirejtshuk & Jarzembowski in Kirejtshuk, 2020 [syn.: Li et al. (2023a)]
- Type species: Notocupes neli Tihelka, Huang & Cai, 2020, by original designation.

References

Primary references

Hong, Y.C. 1983. 北方中侏罗世昆虫化石 [Middle Jurassic Fossil Insects in North China]. Geological Publishing House, Peking. 223 pp. [in Chinese, with English summary] Reference page.
Hong, Y.C. & Wang, W.L. 1990. 莱阳组的昆虫化石 [Fossil insects from the Laiyang Basin, Shandong Province.] Pp. 44–189. In: Regional Geological Surveying Team, Shandong Bureau of Geology and Mineral Resources. 山东莱阳盆地地层古生物 [The Stratigraphy and Palaeontology of Laiyang Basin, Shandong Province.] Geological Publishing House, Beijing. (in Chinese). ISBN 7-116-00667-2. Reference page.
Kirejtshuk, A.G. 2020. Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea. Geosciences 10(2): 73. DOI: 10.3390/geosciences10020073 . Reference page. [treats Notocupes as a synonym of Zygadenia]
Kirejtshuk, A.G., Ponomarenko, A.G., Prokin, A.A., Chang, H., Nikolajev, G.V. & Ren, D. 2010. Current knowledge of Mesozoic Coleoptera from Daohugou and Liaoning (northeast China). Acta Geologica Sinica (English Edition) 84(4): 783–792. DOI: 10.1111/j.1755-6724.2010.00253.x . PDF. Reference page.
Li, Y.-D., Tihelka, E., Newton, A.F., Huang, D.-Y. & Cai, C.-Y. 2023. New species of Notocupes (Coleoptera: Archostemata) from the Middle Jurassic Daohugou beds, with discussion on the generic circumscription. Palaeoentomology 6(4): 398–415. DOI: 10.11646/palaeoentomology.6.4.11 . Reference page. [treats Echinocups as a junior synonym of Notocupes]
Lin, Q-B. 1976. 辽西侏罗系的昆虫化石 [The Jurassic fossil insects from western Liaoning]. Acta Palaeontologica Sinica 15(1): 97–116, 2 pls. Online. [in Chinese, English summary].
Lin, Q.B. 1986. Early Mesozoic fossil insects from South China. Palaeontologia Sinica, Series B 170: 1–112. Reference page.
Ponomarenko, A.G. 1964. [New beetles of the family Cupedidae from the Jurassic of Karatau]. Paleontologicheskii Zhurnal 1964(2): 49–62. (in Russian).
Tan, J.-J., Ren, D. 2006. Ovatocupes: a new cupedid genus (Coleoptera: Archostemata: Cupedidae) from the Jehol Biota (Late Jurassic) of western Liaoning, China. Entomological News 117(2): 223–232. PDF. Reference page.
Tan, J.-J., Ren, D. & Liu, M. 2005. New ommatids from the Late Jurassic of western Liaoning, China (Coleoptera: Archostemata). Insect Science 12(3): 207–216. DOI: 10.1111/j.1005-295X.2005.00026.x . Reference page.

Additional references

Jarzembowski, E.A., Wang, B., Zhang, H. & Fang, Y. 2015 [2014 epub]. Boring beetles are not necessarily dull: New notocupedins (Insecta: Coleoptera) from the Mesozoic of Eurasia and East Gondwana. Cretaceous Research 52(B): 431–439. DOI: 10.1016/j.cretres.2014.03.006 . Reference page.
Jarzembowski, E.A., Zheng, D. & Zhao, X. 2022. Is the beetle Omma (Insecta: Coleoptera) a living fossil? In: Chang, S.-C. & Zheng, D. (eds.) Mesozoic biological events and ecosystems in East Asia. Geological Society, London, Special Publications 521: 161–169. DOI: 10.1144/SP521-2021-56 . ISBN 9781786205834 (electronic), ISBN 9781786205674 (print). Google Books (Preview). Reference page. [treats Notocupes as subgenus of Zygadenia]
Kirejtshuk, A.G., Nel, A. & Kirejtshuk, P.A. 2016. Taxonomy of the reticulate beetles of the subfamily Cupedinae (Coleoptera: Archostemata), with a review of the historical development. Invertebrate Zoology 13(2): 61–190. DOI: 10.15298/invertzool.13.2.01 . Reference page. [mentions Notocupes and Zygadenia; also discusses Chengdecupes as Cupedidae incertae sedis genus]
Lee, S.B., Nam, G.S. & Li, Y.-D. 2022. A new species of Notocupes (Coleoptera: Archostemata) from the Lower Cretaceous (Albian) Jinju Formation in South Korea. Cretaceous Research 140: 105357. DOI: 10.1016/j.cretres.2022.105357 . Reference page.
Li, Y.-D., Tihelka, E., Yamamoto, S., Newton, A.F., Xia, F.-Y., Liu, Y., Huang, D.-Y. & Cai, C.-Y. 2023. Mesozoic Notocupes revealed as the sister group of Cupedidae (Coleoptera: Archostemata). Frontiers in Ecology and Evolution 11: 1015627. DOI: 10.3389/fevo.2023.1015627 . Reference page. [excluded from Ommatidae, temporarily left in Archostemata without familial attribution]
Ponomarenko, A.G. 2000. Beetles of the family Cupedidae from the Lower Cretaceous locality of Semen, Transbaikalia. Paleontological Journal 34(supplementary issue 3): S317–S322. PDF. [treats Notocupes as a synonym of Zygadenia]
Ponomarenko, A.G. 2006. [On the types of Mesozoic archostematan beetles (Insecta, Coleoptera, Archostemata) in the Natural History Museum, London]. Paleontologicheskii Zhurnal 2006(1): 86–94. (in Russian). [English translation: Paleontological Journal 40(1): 90–99. DOI: 10.1134/S0031030106010102 . PDF.] Reference page. [treats Notocupes as a synonym of Zygadenia]
Ponomarenko, A.G. & Ren, D. 2010. First record of Notocupes (Coleoptera: Cupedidae) in locality Daohugou, Middle Jurassic of Inner Mongolia, China. Annales Zoologici 60(2): 169–171. DOI: 10.3161/000345410X516812 . PDF. Reference page. [Zygadenia and Notocupes treated as seperate genera]
Strelnikova, O.D. 2019. Новые купедиды (Insecta: Coleoptera, Cupedidae) из нижнего мела Бурятии [New Cupedids (Insecta: Coleoptera, Cupedidae) from the Lower Cretaceous of Buryatia]. Paleontologicheskii Zhurnal 2019(3): 76–83. (in Russian). [English translation: Paleontological Journal 53(3): 292–299. DOI: 10.1134/S0031030119030146 .] Reference page.
Strelnikova, O.D. & Yan, E.V. 2021. Redescriptions of beetles of the Notocupes generic complex (Coleoptera: Archostemata: Ommatidae) from the Lower Cretaceous of Buryatia. Palaeoentomology 4(5): 499–514. DOI: 10.11646/palaeoentomology.4.5.15 . Plazi. Reference page. [treats Notocupes/Zygadenia and its synonyms as a generic complex]
Strelnikova, O.D. & Yan, E.V. 2023. Redescriptions of the Triassic Notocupes beetles (Archostemata: Ommatidae) from Kyrgyzstan and South Kazakhstan. Palaeoentomology 6(2): 174–190. DOI: 10.11646/palaeoentomology.6.2.9 . Plazi. Reference page. [Zygadenia and Notocupes treated as seperate genera]
Strelnikova, O.D. & Yan, E.V. 2023. On splitting of the genus Notocupes (Coleoptera: Archostemata): new data on morphology and taxonomy. Far Eastern Entomologist 488: 1–28. DOI: 10.25221/fee.488.1 . Reference page. [Notocupes split into four genera: Notocupes, Rhabdocupes, Conexicoxa and Brachilatus]
Tan, J. & Ren, D. 2009. 中国中生代原鞘亚目甲虫化石 [Mesozoic Archostematan Fauna from China]. Science Press, Beijing, 347 pp. ISBN 978-7-03-024607-3. Reference page. [as Ovatocupes and Amblomma]
Tan, J., Ren, D., Shih, C. & Ge, S. 2006. New fossil beetles of the family Ommatidae (Coleoptera: Archostemata) from the Jehol Biota of China. Acta Geologica Sinica (English Edition) 80(4): 474–485. DOI: 10.1111/j.1755-6724.2006.tb00266.x . Reference page.
Wang, W.-l. & Liu, M.-w. 1996. A new species of Notocupes from the Cretaceous of Laiyang Basin, Shandong Province. Memoirs of Beijing Natural History Museum 55: 79–82.

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